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''Physaria filiformis'' (syn. ''Lesquerella filiformis'') is a rare species of flowering plant in the mustard family known by the common names Missouri bladderpod and limestone glade bladderpod. It is native to Missouri and Arkansas in the United States.〔(Biota of North America Program 2014 county distribution map )〕 It was federally listed as an endangered species in 1987 and it was downlisted to threatened status in 2003.〔(''Lesquerella filiformis''. ) The Nature Conservancy.〕 ''Physaria filiformis'' is an annual herb producing several slender, branching stems up to 25 centimeters (10 inches) tall, growing erect or drooping. The leaves vary in shape, and the basal ones reach 2.4 centimeters (0.96 inch) in length. The inflorescence is a raceme of pale yellow flowers with petals half a centimeter to nearly one centimeter (0.4 inch) long. The fruit is a spherical silique.〔(''Physaria filiformis''. ) Flora of North America.〕 The plant is a winter annual, sprouting and forming a basal rosette of leaves in the fall and then producing stems and flower the following spring.〔(''Lesquerella filiformis''. ) Center for Plant Conservation.〕 Seeds are dispersed on the wind and in surface runoff.〔 ''Physaria filiformis'' grows in limestone glades and outcrops of bare bedrock in southwestern Missouri and northwestern Arkansas.〔 The surrounding habitat may be prairie and pasture.〔 Several types of forest may be adjacent to the glades.〔Young, C. C., et al. (2009) (Habitat relationships and management implications for ''Lesquerella filiformis'' Rollins (Missouri bladderpod) on a xeric limestone prairie. ) ''Journal of the Torrey Botanical Society'' 136(2) 233-41.〕 The soil is shallow on the glades and supports few other plants. The plant grows less often in more vegetated areas with deeper soils, probably because other plants successfully compete with it there. It is also less abundant in the shady understory of ''Juniperus virginiana'' forest, as sunlight is a limiting factor in its growth. It can be found in a number of microhabitats, a factor that is important in its management and conservation.〔 Wildfire is important in the maintenance of the limestone glade habitat and even more so its surrounding ecotones. Fire is the mechanism that keeps these habitats open and free of large and woody vegetation that shade out small annuals.〔 Fire prevents ecological succession in the open habitat where the plant grows. Other plants in the local habitat include ''Arenaria patula'', ''Camassia scilloides'', ''Nothoscordum bivalve'', ''Opuntia humifusa'', ''Satureja arkansana'', ''Tradescantia tharpii'', and ''Verbena canadensis''.〔 As the health of the local ecosystem depends on fire, the practice of fire suppression has caused the degradation of the habitat. When fire is prevented, woody vegetation moves in, covering the glades and blocking the light from reaching this small plant.〔 This encroachment of surrounding vegetation is a main threat to the species.〔 Other threats include off-road vehicles, limestone mining, and urban development.〔 Invasive species of plants have been introduced to the habitat, including ''Bromus tectorum'', a competitive grass. The plant may be affected by grazing, trampling, and other disturbance, but these threats are mild because the plant can tolerate light disturbance.〔 When ''Physaria filiformis'' was designated an endangered species there were nine populations.〔USFWS. (Determination of endangered status for ''Lesquerella filiformis'' (Missouri Bladder-pod). ) ''Federal Register'' January 8, 1987.〕 Extensive surveys located more occurrences and many of these were put under the protection of a number of agencies, including the National Park Service and The Nature Conservancy. By 2003 there were about 60 populations and the outlook for the plant had improved enough to warrant its downlisting to threatened status.〔 Most occurrences of the plant are in southwestern Missouri. A large population can be found at Wilson's Creek National Battlefield.〔Young, C. C., et al. (2008). (Monitoring ''Lesquerella filiformis'' Rollins (Missouri bladderpod): Application and evaluation of a grid-based survey approach. ) ''Natural Areas Journal'' 28 370-78.〕 Conservation activities include protection of the habitat in public areas, prescribed burns to replace the natural fire regime, and reduced use of herbicides.〔 ==References== 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Physaria filiformis」の詳細全文を読む スポンサード リンク
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